Khulan tend to walk and feed in a stop-and-go fashion, and regularly lift their heads to look around or make larger uninterrupted movements to the next forage patch. Consequently, distinguishing between walking or standing, standing or feeding, and walking or feeding from values averaged over 4.8-minute intervals must be expected to include substantial noise, as has been shown for other ungulates [50]. Using camera collars with video in combination with raw acceleration data can be expected to improve the ability to also detect short, but costly behaviour like vigilance [18].
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